Uranus is the angel-like, but extremely power-hungry second-in-command of the Secret Society, a criminal organization. Though she served the group's leader, Master, with no question, Uranus had sought to overthrow her and become the new leader of the organization. Prior to joining the Secret Society, Uranus was an ordinary woman and the girlfriend of Duke B. Rambert.
When she learned that Gaia, one of her co-workers, had hosted the first Toshinden tournament behind the Secret Society's back in order to start a rebellion, she declared him a traitor and marked him for death. However, Gaia escaped and went into hiding. Uranus had then sent him an invitation to the second tournament in an attempt to lure him out. Although Gaia had indeed entered, Uranus was eventually killed, along with Master, by him and the rest of the fighters he had aligned himself with during the tournament.
During the events of the fourth tournament, a girl named Puella Marionette carries Uranus' bow and it is strongly implied that Uranus was her mother.