As a Chasseur, Maria has been subjected to the bias that Vampires as a whole are evil, and has thus dedicated her life to eradicating them in the name of her faith. Recently, Maria has been assigned to operate under Roland Fortis, who is new to the Parisian branch of the Chasseurs. When Maria, alongside her colleague, Georges, finds Roland berated within the catacombs beneath Notre Dame, the three work together in order to hunt Vanitas and his vampiric accomplice, Noé Archiviste.

Unexpectedly though, the two factions come to see eye-to-eye and agree to work together in order to seek out Moreau, who has been kidnapping Vampires for his experiments.